Hello, 
I have data of quantitative gentics composed of various traits measurement for multiple population and multiple species. 
In each species one can find few populations and in each population a few traits are measured, so I initially constructed a glmer model as follows: 

model<-glmer(response~fixed effetcs+ (1|Species/Population/Trait) , data=a) 

I needed then to add phylogeny to my analysis and decided to use MCMCglmm. I realised I was unable to take into account the nested structure of the three random effects: 
Species, Population and Trait 
If anyone has an idea of how it should be done, I would be very grateful for help. 

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