[R] Nesting random effects in lmer

[MRak]

Dear R experts,

Could you please tell me how to nest random effects in an lmer model?

Right now, my function looks like this:
Full<-lmer(Consortship~((1|Male)+(1|Female))+MaleRank+MaleAge+MMGroom+MMAggrRankControl+FemaleRank+FemaleAge+MFAggrMating+GroomBirth07+MatNoConsortGroomF,
family=poisson)

However, a) I'm not sure if placing the two random effects (Male and Female)
in parentheses actually achieves nesting, and b) I get an error message
about variable MMGroom (Error in model.frame.default(formula = Consortship ~
((1 + Male) + (1 +  :
  invalid type (S4) for variable 'MMGroom'

Here are a few lines of my data set (unfortunately, my table has 12 columns
and doesn't fit well):

Male MaleRank MaleAge MMGroom MMAggrRankControl Female FemaleRank FemaleAge
MFAggrMating GroomBirth07 MatNoConsortGroomF Consortship
03D 0.756097561 13.48 0 0 15O 0.663157895 8.413415469 0.109289617 0 0 0
03D 0.756097561 13.48 0 0 22S 0.926315789 6.491444216 0.109289617 0 0 0
03D 0.756097561 13.48 0 0 35B 0.442105263 15.36481862 0 0 0.218579235 0
03D 0.756097561 13.48 0 0 38E 0.98 12.35318275 0 0 0.43715847 0.218579235
13I 0.49 10.15 0 0.022537751 25Z 0.27 4.410677618 0.383141762 0 0 0
13I 0.49 10.15 0 0.022537751 35F 0.21 12.3559206 0 1.53256705 0
14A 0.512195122 16.23 0 0 20R 0.59375 6.349075975 0.225225225 0 0

Also is it possible to calculate the RSS of GLMMs?  If so, how can I do
that?

Thank you for your help,

Maria 

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